During the colonial period, the chief source of immigrants to Georgia was England; other important national groups were Germans, Scots, and Scotch-Irish. The number of African slaves increased from 1,000 in 1752 to nearly 20,000 in 1776. After the Revolution, a large number of Virginians came to Georgia, as well as lesser numbers of French refugees from Hispaniola and immigrants from Ireland and Germany. Following the Civil War, there was some immigration from Italy, Russia, and Greece. The greatest population shifts during the 20th century have been from country to town and, after World War I, of black Georgians to northern cities. Georgia suffered a net loss through migration of 502,000 from 1940 to 1960, but enjoyed a net gain of 329,000 during 1970–80 and about 500,000 during 1980–90. From 1985 to 1990, Georgia's net gain through migration was greater than that of any other state except California and Florida. There were net gains of 598,000 in domestic migration and 90,000 in international migration between 1990 and 1998. From 1980 to 1990, the proportion of native-born residents in Georgia fell from 71% to 64.5%. In 1998 Georgia admitted 10,445 immigrants from foreign countries. Between 1990 and 1998, the state's overall population increased 18%. In the period 1995–2000, 965,558 people moved into the state and 624,853 moved out, for a net gain of 340,705, 13,926 of whom were age 65 or over.